X

total mass of dry biomass in the bed (kg-dry-biomass)

Y -s*-m

maximum possible mass of dry biomass in the bed (kg-dry-biomass)

Xo

initial mass of dry biomass in the bed (kg-dry-biomass)

Yq

metabolic heat yield coefficient (J kg-biomass-1)

YW

metabolic water yield coefficient (kg-H2O kg-biomass-1)

YXS

biomass yield from dry substrate (kg-biomass kg-dry-substrate-1)

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